2. Naming the New Genus |
In 1986, nearly a year after our Father's Day excavation, I was awarded a grant from the Smithsonian Institution. Its "short-term visitor" program encourages scientists to study the Smithsonian collections. At the Smithsonian I studied the sauropod dinosaurs and accumulated critical measurements for comparison of Diplodocus and other dinosaurs with the bones of Sam. I extended the travel on that grant to include the American Museum of Natural History (New York City) and the Carnegie Museum of Natural History (Pittsburgh), where the most important specimens of the sauropods Diplodocus, Apatosaurus, and Barosaurus are on display and in study collections.
The information I gathered on this extended travel proved instrumental in my determining the uniqueness of the new bones from New Mexico, and it gave me a firm basis for making size comparisons. According to all comparisons I could make with Diplodocus and Apatosaurus, Sam would be considerably longer, and probably larger in all dimensions. I thus knew we faced a colossal excavation if the bone fragment indeed marked the position of the tail and would lead to the remainder of the skeleton.
Among dinosaurs, sauropods were both the largest and the most conservative. With long necks, long tails, capacious rib cages, and legs like pillars, sauropods changed little physically from the time of their first appearance more than two hundred million years ago (early Jurassic) until their extinction, along with all the other dinosaurs, sixty-five million years ago (end of Cretaceous). They were always enormous and they were all plant eaters. A 1990 summary by J. S. McIntosh lists most of the currently accepted genera of sauropods, along with their ages, principal localities, and comments concerning distinguishing features. (Seismosaurus was not included in that list because its formal description had not then been published.)
Sauropods like Sam flourished in the Jurassic and reached their zenith in diversity at the end of that period, roughly one hundred fifty million years ago. Sam was one of many kinds of sauropods that lived in the late Jurassic. Skeletons of late Jurassic sauropods occur in (relative) profusion in western North America, principally in the Morrison Formation, and in eastern Africa in the Tendagaru beds of Tanzania. The Jurassic Period could well be called the Age of Sauropods.
Sauropods never recovered from a decline that began in the early Cretaceous when angiosperms (flowering plants) began to replace the conifers, cycads, and ferns that had dominated earlier landscapes. Perhaps sauropods survived in the ever-dwindling archaic habitats, while other herbivores with adaptations better suited to feeding on angiosperms replaced them as the dominant plant-eaters of the Cretaceous. Their ultimate disappearance at the end of the Cretaceous scarcely affected the course of history, for their abundance and diversity had diminished so severely that their scant contribution to the dinosaur world at the time of the great extinction could be easily overlooked.
More than 90 genera and 150 species of sauropods have been named, but only a few are relatively well known scientifically. Typical among the Jurassic sauropods was the genus Camarasaurus , a stout animal with relatively short neck and tail. Apatosaurus (as Brontosaurus has been renamed) and Diplodocus belong in another family, differing in their exceptionally long necks and tails and more delicate skulls. All these giants lived in the western United States, likely roaming in herds and feeding constantly on conifers, cycads, and ferns. The weight of most sauropods typically approached twenty tons, with the heavily built Apatosaurus weighing considerably more. Barosaurus, a close relative of Apatosaurus and Diplodocus, had an exceedingly long tail, sometimes described as a whiplash.
The type locality of Seismosaurus in the Ojito Wilderness Study Area northwest of Albuquerque, New Mexico, and the Dry Mesa Quarry southeast of Delta, Colorado--the type locality of Ultrasaurus macintoshi, Supersaurus viviani, and Dystylosaurus edwini.
Mounted skeletons of sauropods now grace the halls of museums around
the world. Increasingly, these skeletons are replicas made from original
reconstructions. And original reconstructions themselves are not the real
thing; they are most often a potpourri, a single skeleton made from
isolated bones--often from several individuals. Reconstructions of
skeletons are not the same as restorations. These two terms have very
different meanings. For example, John Harris distinguishes them in this
way: "the term reconstruction is used in the sense of
piecing together the original but often fragmentary fossilized parts of
extinct animals, whereas restoration is used to describe the
depiction of their original appearance--muscles, flesh, skin, and all."
Thus, a reconstruction is a skeleton, and a restoration is the animal in
the flesh, as though living. Animated restorations, which depict
dinosaurs in their flesh-and-blood glory with movements controlled by
computers, draw huge crowds. Artists' paintings, meanwhile, have long
re-created the living animals in their habitats--giving them a lifelike
quality that is sometimes so realistic that they seem to
be photographs taken by a camera loaded with fast, color-saturated film.
But the real nuts-and-bolts of the dinosaur world is skeletons and the
sites from which the skeletons have been recovered. The real natural
history of dinosaurs resides there, in the bones and the collecting
localities, the only sources for the raw material of dinosaur studies.
The Smithsonian grant gave me the opportunity to study the real bones of
some of Sam's relatives. This award was pivotal in the project, and it
led to the other, larger grants that made excavation beyond the discovery
stage possible.
Before I went to the Smithsonian, I thought Sam was a member of the
genus Diplodocus, a well-known and widespread dinosaur in the Morrison
Formation of Utah, Colorado, and Wyoming. But I had trouble matching the
eight tail vertebrae we had collected during our initial excavation to
known tails of Diplodocus. The proportions were off, the dimensions were
too large, and the anatomical details were different enough to question
the identification of Sam as Diplodocus. The possibility surfaced: maybe
it's a new dinosaur. The fact that the site is hundreds of miles from
other Diplodocus localities in the Morrison Formation added to the
uncertainty.
Eventually I came to believe that Sam could not be Diplodocus or kin
such as Barosaurus or Apatosaurus. The tail bones of Apatosaurus do not
have a deep concavity on their undersurface. The tail bones of Barosaurus
have the deep concavity, like Sam's, but the vertebrae are relatively
short. The closest resemblance was to Diplodocus, but the differences
were still too great. Notably, Sam's vertebrae are proportionally longer
and taller, and the dorsal spines are nearly erect, quite in contrast to
the tail vertebrae of Diplodocus. When I consulted with other sauropod
specialists, they were unable to offer any new information or
interpretations that I had not already considered. I thus concluded that
Sam belonged to a hitherto unrecognized species of dinosaur.
When the New Mexico Museum of Natural History decided to announce the
existence of this new and impressive dinosaur to the public later in
1986, I faced a dilemma. We needed a name for the skeleton, and I could
not assign it a name based on any known dinosaur.
The paleontologist who first describes a fossil as a new species has
the singular responsibility and honor of selecting the name. By
international convention, the name should be latinized according to a universal standard adopted by all
zoologists, The International Code of Zoological
Nomenclature. Its contents read like a legal document, and
issues related to naming of animals resemble court cases.
The choice of name--the technical name--for a dinosaur, or any fossil
organism, is as important to a paleontologist as naming a newborn is to a
parent. We cannot take the matter lightly, because the technical name
will stay with the species forever. Sam and all Sam's kind would be known
by the technical name. The name should have meaning; ideally it should
also be easy to remember and pronounce. And it should be constructed
using the agreed-upon rules designed just for this purpose. In some ways
the honor of coining the name is the most pleasant of our
responsibilities.
That summer, before the press conference organized by our museum
staff, I lay awake at night in my cabin in the mountains east of
Albuquerque, deliberating. I pored over my dictionary of scientific
names, seeking an appropriate root to combine with -saurus
in keeping with the tradition of Apatosaurus, Barosaurus, Camarasaurus,
and dozens of other dinosaurs named under this convention.
After several weeks of searching for a name, I made a decision. I
chose Seismosaurus. Seismo is the Latin root for "shaking."
It is familiar in words like seismic and
seismology , all relating to ground-shaking generated by
earthquakes or underground blasts. Sam would be the earth-shaking
dinosaur. I searched the technical literature to ensure that this name
had not already been taken; if so, the name would have been "preoccupied"
and not available for any newly discovered species.
I was lucky. Seismosaurus hadn't been used before; my first choice was
available. As long as I didn't attach a species name to this informal
genus, any publication of the name in print would be safe from technical
nullification--provided I was right that Sam was a new genus. I would use
this name in the press conference.
Floodlights blinded me, and the array of microphones seemed like
menacing tentacles of a giant octopus. Paleontologists are not trained in
graduate school for press conferences. I briefly explained what we had
found, unveiled the bones, and revealed the new informal name. I reported
that this individual of Seismosaurus was probably at least 110 feet long,
comparing it with our own mounted skeleton of Camarasaurus (which is only
about 50 feet long.) This put Sam in the ranks of the
supergiants, not the giants.
I then demonstrated how I made that calculation by direct proportions
with Sam's closest relative, Diplodocus. Each vertebra was at least 20
percent longer than corresponding vertebrae in Diplodocus, and with the
disproportionately tall neural spines, there was the possibility that the
overall length of the tail (and indeed the entire body) was also
disproportionately long. Although I had data in hand that indicated a
more likely length of 120 feet or more, I chose to conservatively
estimate Sam's length at 110 feet--which is longer by 23 feet than the
longest specimen of Diplodocus, the previously accepted longest dinosaur.
(Later I would revise these figures upward.)
Questions came from all directions as each reporter sought a different
angle to develop. Sam's new name, Seismosaurus, caught on immediately,
capturing the reporters' imaginations. One reporter asked rhetorically
why I didn't select a name like Superdoopersaurus to follow the recently
invented names Supersaurus and Ultrasaurus for two supergiant dinosaurs
that were discovered in Colorado. That quip lightened up the discussion.
We had prepared an exhibit case for the four vertebrae, set beneath
the Camarasaurus skeleton for comparison with its tail
vertebrae. Comparison with a skeleton of Diplodocus or Apatosaurus would
have been more appropriate, since Seismosaurus belongs in the same family
with these two familiar dinosaurs, but the Camarasaurus skeleton was the
closest comparison we could make with the exhibits available. The
corresponding vertebrae in the tail of the Camarasaurus skeleton are
ridiculously small by comparison. The display emphasized the
extraordinary size of the new dinosaur, even though we had only four
bones to present to the public.
The press conference generated a surge of media attention, more than I
ever imagined. Sam (rather, Seismosaurus) was spectacular.
My allusion to "earth-shaking" proved ironic, for the next year we
would initiate experiments in remote sensing to look for more of Sam's
bones hidden deeper in the mesa. Artificially generated sound waves (from
a fancy shotgun) would help us "see" bones without digging. That
technology was called seismic tomography.
To formalize the name Seismosaurus I needed to fully describe the
eight tail vertebrae on which the determination had been made. This
description had to be published in a technical journal. I had to
distinguish the bones from all other dinosaurs, including Sam's closest
relatives, Diplodocus, Apatosaurus, and Barosaurus. This might appear to
be a dry and simple task, but technical descriptions of new species are
difficult and demanding. Putting into words the description of a bone or
skeleton is an extraordinary challenge of communication in the use of our
wonderfully versatile language. To succinctly describe an object that is
as irregular as a bone, and to do so in words that others can understand
without ever seeing it, is immensely satisfying. I, of course, had a
science illustrator draw the bones from various perspectives, but in a
scientific journal the words are definitive.
Formal description would not be done overnight. What is more, I knew
that if I waited I would almost surely have more bones upon which to base
the genus-making description. Publication would therefore wait.
Seismosaurus was not, however, a complete name for formal publication.
Living and extinct organisms are given binomial names. The principle of
binomial nomenclature is that every organism is given a pair of names: a
genus name that is capitalized and a species name that is not. Humans
belong to the genus Homo. We share this genus with no living species but
with several extinct hominid species--Homo erectus , for
example. Our species is sapiens. We are therefore Homo
sapiens --presumably, "wise" humans.
The informal name Seismosaurus established at the press conference could not be formalized without a species name
to follow it. That left me with another decision: what to call Sam's
species in the formal description.
Sometimes a species name is coined for an anatomical feature, or for a
locality, or for a person, such as the discoverer or a patron of the
project. Several times I half-jokingly offered to name the species after
anyone willing to donate half a million dollars to the project, and the
genus after anyone willing to donate a million dollars. My circle of
friends is not wealthy; I got no takers.
I considered naming the species for Arthur Loy and Jan Cummings, who
together found the bones. In fairness, however, I recognized not only
Arthur and Jan as the discoverers but also their friends Frank Walker,
who brought the bones to my attention and showed them to me, and Bill
Norlander, the fourth member of this fraternity of hiking buddies. I
couldn't name the species for all four, and naming it for one wouldn't be
fair to the others.
What about geography? The correct latinization of ojito ,
for the site in the Ojito Wilderness Study Area, would be
ojitoensis . The pronunciation would thus be a puzzle to
everyone not familiar with Spanish etymology, and I dislike
tongue-twister names anyway.
What about anatomy? Most of Sam's anatomical features are subtle and I
couldn't find any one trait in particular that would by itself
characterize the species. I thought about referring to the size of the
new species by using longus or colossus , but
these names didn't seem appropriate either.
I settled on naming the species for the Reverend James Hall, director
of the Ghost Ranch Conference Center, and his wife Ruth Hall, an amateur
paleontologist who inspired several professional careers by her teaching.
Ghost Ranch is in northern New Mexico, a study center owned by the
Presbyterian Church in the canyonlands north of Santa Fe made famous by
the artist Georgia O'Keeffe. On its 23,000 acres is one of the richest
and most spectacular dinosaur sites in the world, a quarry where at least
a thousand individuals of the little predatory dinosaur Coelophysis have
been excavated. Jim and Ruth together supported paleontology in and around Ghost Ranch and northern New
Mexico for a quarter century. I began working there in 1985 and continue
with several active projects related to the Coelophysis quarry. Ghost
Ranch has since established the Ruth Hall Museum of Paleontology,
organized by Lynett Gillette, the museum's first curator.
Seismosaurus would thus bear the simple species name "halli."
Seismosaurus halli , or "Hall's earth-shaker dinosaur," it
would be.
I now had the name necessary for publication, and soon I had more
bones. I submitted my description to the Journal of Vertebrate
Paleontology in 1989. The review process stretched on, however.
Scientists are naturally skeptical of claims of new species (and, even
more so, genera), and the peer reviewers of my paper took their task
seriously. I had to respond to their criticisms; producing acceptable
revisions added another year to the publication date. The descriptive
paper was finally published in 1991. Prior to that time, I had given a
talk at a scientific symposium (1986) and had published a short abstract
(1987), referring to Sam as "a giant sauropod" or "a new giant sauropod"
from the Morrison Formation of New Mexico. I had also written a popular
article on the excavation for the Ghost Ranch Journal . But
publication of a formal description and a full scientific name in the
Journal of Vertebrate Paleontology made it official.
To paleontologists the full and correct name for the new species is
now "Seismosaurus hallorum Gillette 1991." My initial name
proved to have an incorrect Latin ending of a genitive singular--a
mistake recognized by George Olshevsky, a dinosaur classification
aficionado. So it was changed to the plural form. I formally assigned it
to the family Diplodocidae, the family that includes the giants
Diplodocus, Barosaurus, and Apatosaurus and the supergiant Supersaurus,
all from the Morrison Formation of western North America.
From my original coining of Seismosaurus to technical publication of
the name Seismosaurus halli (more properly Seismosaurus
hallorum ) took five long years. With the formal publication of the
name and description of Sam's bones as the basis for the
new species, our initial goals had been achieved: we had defined the
species, identified its distinguishing characteristics, established the
geologic age in which the animal lived (late Jurassic) and the geographic
position of the site (the southern end of the Morrison Formation), and
presented the data and interpretive calculations that would verify Sam's
size--then calculated as between 128 feet and 170 feet, or between 39 and
52 meters--in the technical literature, through the rigors of peer
review.
Sam's position in the scheme of classification of animals can be
succinctly summarized. The fundamental unit of classification is the
species. Taxonomic categories above the species level are increasingly
subjective, generally arranged in hierarchical order. For Sam, the full
classification using traditional ranks is as follows:
Phylum Cordata
Some paleontologists prefer to separate dinosaurs from the reptiles
into a distinct class: Dinosauria. Usually by that convention, Dinosauria
includes only dinosaurs, but some paleontologists place birds in the same
class, subsuming the traditional class Aves into Dinosauria. At issue are
the questions of origins and the philosophy of establishing these
evolutionary hierarchies. In recent years applications of the principles
of cladistics, and with them a new system of nomenclature, have clarified
many questions of ancestry, but the basic unit of classification, the
binomial (genus and species) remains largely unaffected.
This naming, this classification, this identification and formal
description of Sam depended ultimately on the bones. How does one find
more bones hidden, and perhaps scattered, inside a mesa?
Skeletal reconstruction of Diplodocus
carnegii, probably the closest known relative of
Seismosaurus. This slender sauropod is known from
essentially complete skeletons from the Morrison Formation. Its skeletal
anatomy is well established, and it formed the basis for most comparisons
with Sam's skeleton.
Muscular anatomy of Diplodocus
carnegii . This kind of drawing is a prerequisite for producing
a reasonable in-the-flesh restoration. Seismosaurus was similar, but had
more massive hips, stouter (but not longer) legs, and a tail that
differed in some important ways.
First display of the tail vertebrae at
the New Mexico Museum of Natural History, Albuquerque. This vertebra is
no. 20, counting from the base of the tail. It was the anterior-most of
the original eight excavated in 1985.
A partially prepared Seismosaurus caudal
vertebra. The sandstone in which this bone was encased was so perfectly
matched in color and texture that distinguishing bone under ordinary
lighting conditions (such as photographed here) was almost
impossible.
Subphylum Vertebrata (all animals with backbones)
Class Reptilia (all reptiles including dinosaurs)
Order Saurischia (the giant quadrupedal herbivorous dinosaurs and
bipedal carnivores)
Suborder Sauropoda (the giant quadrupedal long-necked herbivores)
Family Diplodocidae (relatives of Diplodocus)
Genus Seismosaurus
Species hallorum, correctly expressed as the binomen
Seismosaurus hallorum.